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After Selene helps take the bullet out of David, they are found by two vampires who escort them back to the sanctuary, where Selene is introduced to the recruits and is told she will teach them how to fight against lycans. Thomas speaks before Cassius and the other vampire council members to convince them to allow Selene to train new death dealer recruits. He lays out a plan of attack so that the lycans can destroy the vampire's sanctuary, but Marius also wants Eve's blood to gain a great power. Marius meets with his group in their hideout. Semira also hoped to succeed Viktor as an elder after Selene killed him, and she, along with a vampire leader named Cassius (James Faulkner), wants to prosecute her for it. They discuss the growth of Marius's lycan army and the destruction of the Western coven, with the main concern being to prevent a potential full-scale lycan attack. Varga brings Thomas to Semira (Lara Pulver), a high-ranking vampire in the coven. David's father Thomas (Charles Dance) arrives at the home of the Eastern vampire coven, where he is greeted by Varga (Bradley James). He shoots one of the lycans in the foot as an incentive to send a message to Marius, but David also ends up getting shot. She battles the men as they morph into their wolf forms, just as David (Theo James) arrives to help her.
Selene rides her motorcycle as she's being pursued by a group of lycans. The lycans have a new leader named Marius (Tobias Menzies). Without Michael, Eve, or her coven, Selene feels alone as both vampires and lycans are hunting her. Selene was forced to keep Eve in hiding to keep her safe as she went to look for Michael. Selene and Michael's daughter Eve (India Eisley) is the first pureblood hybrid. She was betrayed by the vampire elder Viktor (Bill Nighy) and later met and fell in love with Michael Corvin (Scott Speedman), who would become the first vampire-lycan hybrid. The vampire Selene (Kate Beckinsale) has been a death dealer for thousands of years in the war against the lycans (werewolves). The film starts with a recap of the previous 'Underworld' films.The synopsis below may give away important plot points. And it's while they are there at the stronghold that David learns the truth about his lineage. Eventually Marius learns of this and sets out to find Selene and find out for sure if she knows where her daughter is. Eventually David learns of what Semira is doing and tries to get Selene away and takes her way to the only other Vampire stronghold there is. She then reveals she wants Selene for her blood. Semira then has her man kill the trainees with Selene's weapon.
It's when she battles one of Semira's people, that he subdues her with a toxin that paralyzes her. They grant it and send word to Selene who comes and trains the aspiring Death Dealers. At the same time, Semira, Vampire Elder implores Thomas, another elder and father of David, Selene's ally to convince the Council of Elders to give Selene whom they deem a traitor, amnesty so that she could train new Death Dealers to deal with the Lycans. But when they attempt to capture her, they fail and she tells one of them to tell Marius, she doesn't know where her daughter is her daughter has cut ties with her. He thinks if he can get the hybrid daughter of Vampire Death Dealer Selene, her blood will be the key to defeating the Vampires once and for all.
The war between the Vampires and the Lycans wages on with the Lycans led by Marius are close to defeating the Vampires.
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